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The River Rother runs for 35 miles from Rotherfield, East Sussex, entering Kent after Bodiam where it follows the county border to Wittersham, where it rejoins East Sussex to the mouth at Rye.

There are two other River Rothers in Yorkshire and in Hampshire to West Sussex.

The Rother Levels are an area of reclaimed land around the lower course of the River Rother on the Kent - East Sussex border. At one time the sea occupied the area, but gradually the area silted up and over time was drained and reclaimed. The Isle of Oxney sits in the middle of the levels.