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The Cuckoo Line is a now disused railway line which linked Polegate and Eridge in East Sussex, from 1880 to 1968. The railway line served the following Sussex communities: Polegate, Hailsham, Hellingly, Horam for Waldron, Heathfield, Mayfield, Rotherfield and Eridge. Services continued through Eridge and onward via Groombridge to Tunbridge Wells.
The Hailsham-Eridge section closed in 1965, the Polegate-Hailsham branch surviving until 1968. Eridge-Tunbridge Wells closed in 1985, and this line has been resurrected as the Spa Valley Railway.
The section of the line from Polegate to Heathfield is now National Cycle Network Route 21 and The Cuckoo Trail.
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Route 21 of The National Cycle Network runs south from Greenwich out of London through Lewisham (the Waterlink Way) to Crawley, and then via East Grinstead and Eridge to Heathfield and Eastbourne, a distance of 95 miles.
The section between Groombridge and East Grinstead follows the Forest Way. Link
See Link for a map of the route.

This 13 mile (21km) surfaced path through the Sussex countryside is mostly traffic free and is a safe route for walkers and cyclists of all ages, forming part of the National Cycle Network – route 21.

The picturesque trail follows the former 'Cuckoo Line' railway track and stretches from Heathfield to Eastbourne Park. It passes through Horam, Hailsham and Polegate.
Link